Mass flow theory solution is forced to move through phloem more sugar continues to be loaded phloem is under positive pressure an active sink is required for transport to continue. The pressure flow hypothesis helps explain how dissolved sugars move from sugar sources to sugar sinks. This creates turgor pressure, also known as hydrostatic pressure, in. The flow stops, when turgor pressure both in a and b are equal. Discussion and essay questionsset 3 photosynthetic. This sugar is then moved into sieve tube cells by active transport. Before moving into the source cells present in the phloem, the prepared food is converted into sucrose. The models assume a continuous active loading and unloading of translocate but differ in the site of loading and unloading and the route of water to the sieve tube. Five key facts and their discussion can lead to the conclusion that a turgor pressure difference along sieve tubes of the phloem drives a mass flow. However if a high turgor pressure is maintained at a, there is a continuous flow from a to b this is possible in the plant by the continuous synthesis of food on reaching chamber b the water will flow into the outer vessel xylem. At present, the most widely accepted theory for movement of substances in the phloem is called the pressure flow or mass flow hypothesis.
The following are the answers to the practice questions. Now, one theory is generally accepted as the correct explanation for translocation. The mass flow hypothesis can be explained as the mass flow of solute taking place from the source that is the mesophyll cells where the solutes are produced. Proposed by munch in 1926, the pressureflow hypothesis states that a mass flow of water and sugar is driven by a pressure difference through sieve tubes of. The monrokellie doctrine was first described by dr alexander monro and dr george kellie.
Also, the very high turgor in the phloem of herbs suggests that pressure has functions beyond that of sap conduction. The pressure flow hypothesis, also known as the mass flow hypothesis, is the bestsupported theory to explain the movement of sap through the phloem. Water may merely trickle along a creek bed in a dry season, but rush quickly and under great pressure after a heavy rain. This flow occurs along the gradient of the turgor pressure from a region of higher solute concentration to a. Munch explained the translocation of the organic food material through the phloem tissue. Among them, munchs 1930 hypothesis is most convincing. A than the other b, are connected by a tube c and dipped in water. Mechanism of translocation munch pressure flow hypothesis sugar loaded into phloem increase in solute potential leads to increase in water uptake from xylem builds up hydrostatic pressure unloaded at sink. A mathematical treatment of munchs pressureflow hypothesis. Pdf revisiting the mnch pressureflow hypothesis for long. Mass flow hypothesis was proposed by german scientist ernst munch in 1930, who described the movement of sap via phloem. Pressure flow hypothesis article about pressure flow. The theory behind mass flow hypothesis which is also called as pressure flow hypothesis describes the movement of sap via phloem, proposed by the german. A high concentration of organic substances, particularly sugar, inside cells of the phloem at a source, such as a leaf, creates a diffusion gradient that draws water into the cells from the adjacent xylem.
Process control pressure, flow, and level processes courseware sample 87996f0. The pressure flow hypothesis, also known as the mass flow hypothesis, is the bestsupported theory to explain the movement of food through the phloem. Munch pressure flow model munch, a german plant physiologist, proposed in 1930, a simple physical model which can be tested in the laboratory for the mechanism of phloem transport. Jun 25, 2015 transport in plants,class 11,chapter 11,biology phloem transport the pressure flow hypothesis in mal duration. Angiosperm organization of the vascular tissue britannica.
While investigators believing in occluded plates favored the electroosmotic theory fensom, 1957. Testing the munch hypothesis of long distance phloem. As glucose is made at the source by photosynthesis for example it is. Similarly, as blood volume decreases, pressure and flow decrease. Explain pressure flow hypothesis of translocation of. Angiosperm angiosperm organization of the vascular tissue. The steady state solutions of two mathematical models are used to evaluate munchs pressureflow hypothesis of phloem translocation. The major steps in the mass flow theory are as follows.
It was proposed by ernst munch, a german plant physiologist in 1930. How would you test the hypothesis in gymnosperms e. What is pressure flow or mass flow hypothesis in transport of. The pressure flow hypothesis introduced by ernst munch in 1930 describes a mechanism of osmotically generated pressure differentials that are supposed to drive the movement of sugars and other solutes in the phloem, but this hypothesis has long faced major challenges. Sugars begin at a source and are pumped into phloem tube cells. This flow occurs along the gradient of the turgor pressure from a region of higher solute concentration to a region of lower solute concentration.
Blood flow, blood pressure, and resistance anatomy and. If another region of the plant needs sugars, they are actively pumped out of. A mathematical treatment of munchs pressureflow hypothesis of phloem. Explain the pressure flow hypothesis by first completing the statements with the terms provided, and then rearranging the statements so that they appear in chronological order. From 1930s to the mid1970s, the mechanism of phloem translocation was a subject of research. In stems, the vascular tissue is organized into many discrete vascular bundles. This paper presents a model based on the munch pressureflow hypothesis constructed from the navierstokes equation for the motion of a viscous fluid and the vant hoff equation for osmotic pressure. The pressure flow hypothesis is still called a hypothesis, it is agreed to have large question marks by it, and many alternative hypotheses have been presented and have received serious attention over the years. Revisiting the munch pressureflow hypothesis for long. Pressure flow hypothesis pdf writer pressure flow hypothesis wikipedia.
In questioning pressure flow, i am not being quite as revolutionary as when i question electron bonding. This flow occurs along the gradient of the turgor pressure. The anatomy of roots and stems is discussed in their. In the roots, the vascular tissue is organized within a single central vascular cylinder. As shown in figure, two osmometers, one containing solute at higher concentration. According to this hypothesis, organic substances move from the region of high osmotic pressure to the region of low osmotic pressure in a mass flow due to the development of a gradient of turgor pressure fig.
Pressure flow hypothesis synonyms, pressure flow hypothesis pronunciation, pressure flow hypothesis translation, english dictionary definition of pressure flow hypothesis. Technical note 12, differential pressure mass flow meter, rev. According to this theory, food substances in solution organic solutes flow from a source, where water enters by osmosis e. The mass flow hypothesis in terms of plants, is the following. The textbook points out that the pressure flow hypothesis has not been confirmed for gymnosperms, which have sieve cells, but not sieve tubes. This means the hydrostatic pressure in phloem increases which causes mass flow. Proximity of source to sink is critical sinks may change during life cycle young leaves net import as mature become exporter roots, shoots, tubers, fruits are strong sink. Chapter 6 design of pe piping systems 158 11 12 where pr pressure rating, psi hds hydrostatic design stress, psi table 11 a f environmental application factor table 12 note. Our data provide strong support for the munch hypothesis as a. While the movement of water through xylem tissue is relatively well understood, the movement of soluteladen sap through phloem is not. Pdf testing the munch hypothesis of long distance phloem.
Find out information about pressure flow hypothesis. Water then follows by osmosis, creating pressure in the tube at the source of the sugars. Pressure flow hypothesis definition of pressure flow. Transport of food substances organic solutes in solution. Spanner, 1958, 1970, the pressure flow hypothesis was supported by the group believing in open pores and that occlusion is due to preparation. Excess sugars can be stored in areas such as roots to be used later. Mass flow hypothesis the mechanism of transportation in plants. According to the pressureflow hypothesis,which of the following is not true. Pdf a mathematical model of the munch pressureflow hypothesis for long. Use the pressure flow hypothesis to explain how sugar can move from the seed into the cells of the plant embryo. Long distance transport in plants occurs in sieve tubes of the phloem. The derivation is for a solution inside a long, narrow, rigid tube with no radial fluxes of solute through the tube walls. Teaching the pressureflow hypothesis of phloem transport in a. As glucose is made at the source by photosynthesis for example it is converted to sucrose a dissacharide.
When sinks need sugar, the pressure difference between the source and sink causes dissolved sugars to move to the area of need. The pressure flow hypothesis introduced by ernst mu nch in 1930 describes a mechanism of osmotically generated pressure. Get a printable copy pdf file of the complete article 1. What is the difference between symplastic and apoplastic unloading of phloem in sink tissue. The movement of water into a nutrient rich region of the phloem decreases the pressure in that region would a year of drought produce a thick or a thin tree ring.
This is a question and answer forum for students, teachers and general visitors for exchanging articles, answers and notes. The relationship between blood volume, blood pressure, and blood flow is intuitively obvious. The theory behind mass flow hypothesis which is also called as pressure flow hypothesis describes the movement of sap via phloem, proposed by the german physiologist ernst munch in 1930. Transport in plants,class 11,chapter 11,biology phloem transport the pressure flow hypothesis in mal duration. The tissue of vascular plants that conducts food produced by photosynthesis to all parts of the plant and consists of. This model of how phloem works is based on the relationship between sinks and sources. As blood volume increases, pressure and flow increase. The hormesis hypothesis article in medical hypotheses 825 may 2014 with 546 reads how we measure reads. Now that weve covered sinks and sources, lets look at the pressure flow hypothesis.
The models assume a continuous active loading and unloading of translocate but. It proposes that water containing food molecules flows under pressure through the phloem. Therefore, water flows by osmosis from adjacent cells. The osmotic pressure of the sieve tubes is decreased.
A high concentration of organic substances, particularly sugar, inside cells of the phloem at a source, such as a leaf, creates a diffusion gradient that draws water into the cells from the. What is pressure flow or mass flow hypothesis in transport. The pressure flow hypothesis, also known as the mass flow hypothesis, is the bestsupported. Active transportactive movement of sugar sucrose at the source into phloem cells causes the water potential of phloem contents to become more negative. Learn vocabulary, terms, and more with flashcards, games, and other study tools. Mass flow hypothesis the mechanism of transportation in. The pressure flow hypothesis the bestsupported theory to explain the movement of food through the phloem is called the pressure flow hypothesis. Starlings hypothesis states that the fluid movement due to filtration across the wall of a capillary is dependent on the balance between the hydrostatic pressure gradient and the oncotic pressure gradient across the capillary. Theory overview of flow measurement using differential.
Sugars are loaded into the sieve tubes of the phloem at the sugar source leaf. The environmental application factors given in table 12 are not to be confused with the design factor, df, used in previous editions of the ppi handbook and in older standards. Vascular tissue is organized into discrete strands called vascular bundles, each containing xylem and phloem. How plants send sugars from sources to sinks dummies. The tissue of vascular plants that conducts food produced by photosynthesis to all parts of the plant and consists of sieve elements, fibers, and. Pressure flow theory the pressure flow theory explains how sap moves in a plant from source to sink. Munch, a german plant physiologist, proposed in 1930, a simple physical model which can be tested in the laboratory for the mechanism of phloem transport. In the end of this link you can have a quick and easy explanation for the mass flow hypothesis. Massflow hypotheses include the pressureflow hypothesis, which states that flow into sieve tubes at source regions places of photosynthesis or mobilization and exportation of storage products raises the osmotic pressure in the sieve tube. Explain pressure flow hypothesis of translocation of sugars.
Water moves from the xylem vessels into the adjacent phloem, thereby increasing the hydrostatic pressure in the phloem. Jun 02, 2016 long distance transport in plants occurs in sieve tubes of the phloem. Various theories have been put forward to explain the mechanism of phloem conduction but they are not fully satisfactory. Translocation of food through phloem definition, examples. The pressure flow hypothesis of food movement video. Active transport moves sugars into the sieve tube from surrounding tissues. A highly concentrated organic sugar especially sugar in the cells of phloem from a source like a leaf forms a diffusion gradient which draws water in the cells from adjacent xylem. The glucose is prepared at the source by the process of photosynthesis and is converted to sucrose sugar. The accepted mechanism needed for the translocation of sugars from source to sink is called the pressure flow hypothesis. It describes the relationship between the contents of the cranium and intracranial pressure. According to the pressure flow hypothesis, food is prepared in the plant leaves in the form of glucose.
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